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On June 20, 1943 one of forty-six aircraft that scrambled intercept Japanese fighters and bomber approaching Darwin (Raid #55). Took off piloted by piloted by F/O A. Hughes then climbed toward the enemy formation at 27,000'. This aircraft made a firing pass at two Ki-49 Helens, then dived on a "Zeke" [Ki-43 Oscar]. Following this attack, the Spitfire's engine seized forcing it to make a glide landing on the beach at Lee Point. Pilot Hughes was unhurt.
On June 29, 1943, despite being partially submerged by the tide, this Spitfire was successfully salvaged by 14 ARD. Repaired, this aircraft was returned to 54 Squadron on March 11, 1944.
On April 7, 1944 during take off from Wyndham Airfield piloted by P/O A. E. Featherstone the engine failed causing the aircraft to crash at the end of the strip and somersaulted, landing on its back. During the crash, Featherstone was slightly injured. Afterwards, transfered to 7 RSU. On May 16, 1944 converted to components and officially written off.
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